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Based upon years of work using hydrolysis to break down and analyze yeast nucleic acids, Levene proposed that nucleic acids were composed of a series of nucleotides, and that each nucleotide was in turn composed of just one of four nitrogen containing bases, a sugar molecule, and a phosphate group. Levene made his initial proposal in 1919, discrediting other suggestions that had been put forth about the structure of nucleic acids. In Levene’s own words, “New facts and new evidence may cause its alteration, but there is no doubt as to the polynucleotide structure of the yeast nucleic acid” (1919)..

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Mercutio is talking about a medlar fruit, which was colloquially referred to as an “open arse,” for reasons that can never be adequately explained. However, there is no such thing as a poperin pear it’s another old timey play on words. Separate “poperin” into its three syllables and you get an Elizabethan penis euphemism “pop ‘er in.”.

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